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MONTANA 



Natural Heritage 

^j^ Program MONTANA An IM A L 







A program of the 

Natural Resource Information System, 

Montana State Library 



Species of Concern july2004 



Introduction 

This report is produced on an annual basis jointly by 
the Montana Natural Heritage Program (MTNHP) and 
Montana Fish Wildlife and Parks (MFWP). Montana 
Species of Concern are native animals breeding in the 
state that are considered to be "at risk" due to declining 
population trends, threats to their habitats, and/or 
restricted distribution. 

Also included in this report are Potential Species of 
Concern -- animals for which current, often limited, 
information suggests potential vulnerability or for which 
additional data are needed before an accurate status 
assessment can be made. Many of these species have 
ranks reflecting the uncertainty of their status, and we 
hope that highlighting them as Potential Species of 
Concern will encourage biologists to conduct surveys and 
submit new data. The category "Species on Review" 
included in the 2003 report has been merged into Potential 
Species of Concern. 

Status determinations are made by MTNHP and 
MFWP biologists in consultation with representatives of 
the Montana Chapter of the Wildlife Society, the Montana 
Chapter of the American Fisheries Society and other 
experts. The process for evaluating and assigning status 
designations uses the Natural Heritage Program ranking 
system, described below, which forms the basis for 
identifying Montana Species of Concern. 

This report identifies 119 vertebrate and 58 inverte- 
brate Species of Concern. Another 37 vertebrates and 50 
invertebrates are identified as Potential Species of 
Concern. See the "Box Scores" below for totals by animal 
group. Thirteen species included in the 2003 report have 
been dropped from consideration (see page 11). 




Coeui d'Alene Salamander 
{Plethodon idahoensis) 

Illustration h\ John Cailsi 



Heritage Program Ranks 

The international network of Natural Heritage Programs 
employs a standardized ranking system to denote global 
and state status. Each species is given a global (G) rank, 
denoting range wide status, and a state (S) rank for its 
status in Montana. Status ranks range from 1 (greatest 
concern) to 5 (least concern). Global ranks are assigned by 
scientists at NatureServe (the international affiliate organi- 
zation for the heritage network) in consultation with 
biologists in the natural heritage programs and other 
taxonomic experts. State ranks are determined jointly by 
MTNHP and MFWP biologists, in consultation with other 
state experts. 

The combination of global and state ranks helps 
describe the proportion of a species' range and/or total 
population occurring in Montana. For instance, a rank of 
G3 S3 indicates that Montana comprises most or a very 
significant portion of an animal's total population. In 
contrast, an animal ranked G5 S 1 often occurs in Montana 
at the periphery of its much larger range; thus, the state 
supports a relatively small portion of its total population. 



Species of Concern 



Amphibians.. 

Birds 

Fish 

Mammals 

Reptiles 

Invertebrates 
Invertebrates 
Invertebrates ■ 



Insects .... 
Mollusks . 
Other 



Box Scores 

Potential Species of Concern 

...5 Amphibians 

,. 60 Birds 18 

.. 18 Fish 10 

.. 27 Mammals 9 

.. 9 Reptiles 

.. 29 Invertebrates - Insects 48 

.. 22 Invertebrates - Mollusks 2 

.. 7 Invertebrates - Other 



What^s on the Lists 

Montana Species of Concern are defined as vertebrate 
animals with a state rank of S 1 , S2, or S3. Vertebrate 
species with a rank indicating uncertainty (SU) or a "range 
rank" extending below the S3 cutoff (e.g., S3S4) are 
considered Potential Species of Concern. 

Because documentation for invertebrates is typically 
far less than for vertebrates, only those ranked SI or S2 are 
included as Species of Concern. Invertebrates ranked S3 
are included only if they are also considered globally 
vulnerable (G3 or higher) or if experts agree their 
occurrence in Montana has been adequately documented. 
All other invertebrates ranked S3 are treated as Potential 
Species of Concern. Also included as Potential Species of 
Concern are globally common (G4 or G5) invertebrates with 
a range rank extending below the S2 cutoff cited above 
(e.g., S2S4). Invertebrates ranked as SU have been 
dropped from Potential Concern. 

How Species are Ranked 

This year, we began implementing a more rigorous 
methodology with more consistent and quantifiable criteria 
to assign status ranks. The goal is to improve the accuracy 
of ranks and better document the basis for each rank. 
Having a rigorous and well-documented ranking procedure 
is especially important to MFWP, which is using status 
ranks to help set priorities for the state's Comprehensive 
Fish and Wildlife Plan. 

This new approach to assessing species' status is 
based on methods developed by NatureServe (Master et al. 
2003). In December 2003, a group of staff and other experts 
met to review the proposed process, tested it on a variety 
of taxa, and endorsed its use in the 2004 status review. 
Staff then contacted Montana experts for each animal 
group and asked them to rank species according to six 
separate criteria. These criteria included population size, 
area of occupancy in Montana, short and long-term trends, 
threats, inherent vulnerability, and specificity to 
environment. Based on this expert input, a preliminary rank 
was calculated, which then was reviewed with key experts. 

It is important to note that while the ranking process 
has changed, the definitions and the fundamental criteria 
remain the same. The principle difference is that criteria are 
now being applied in a more rigorous and consistent 
manner. Detailed documentation of the criteria and 
assessment process are available on the MTNHP website 
at mtnhp.org/animals/index.html. 

This status assessment process was completed for all 
of Montana's amphibian, reptile and mammal species. Due 
to the large number of bird species in the state, only those 
already on the 2003 Species of Concern list were evaluated 
to date; the remaining bird species will be assessed in 
future revisions. Updated ranks for fish are not included in 
this report, to allow the Montana Chapter of the American 
Fisheries Society additional time for review; they will be 
reported in the 2005 update. The new ranking methodology 



has not yet been applied to invertebrates, so the ranks 
presented in this report reflect the traditional ranking 
approach. 

Definitions of Ranks used in this Report 

Gl / S 1 At high risk because of extremely limited and/or 
rapidly declining numbers, range, and/or habitat, 
making it highly vulnerable to global extinction or 
extirpation in the state. 

G2 / S2 At risk because of very limited and/or declining 
numbers, range, and/or habitat, making it 
vulnerable to global extinction or extirpation in the 
state. 

G3 / S3 Potentially at risk because of limited and/or 

declining numbers, range, and/or habitat, even 
though it may be abundant in some areas. 

G4 / S4 Uncommon but not rare (although it may be rare in 
parts of its range), and usually widespread. 
Apparently not vulnerable in most of its range, but 
possibly cause for long-term concern. 

G5 / S5 Common, widespread, and abundant (although it 
may be rare in parts of its range). Not vulnerable in 
I I most of its range. 

GU/ SU Currently unrankable due to lack of information or 
due to substantially conflicting information about 
status or trends. 

GH/SH Historically occurred; may be rediscovered. 

GX/SX Believed to be extinct; historical records only. 

Other codes and rank modifiers: 

B / N State rank modifiers indicating the breeding status 
for a migratory species; B = Breeding, N = Non- 
breeding. 

HYB A global rank denoting a hybrid. 

M A state rank modifier indicating migratory 

stopover status for a species. 

Q A global rank modifier indicating that there are 

taxonomic questions or problems. 

T Denotes the rank for a sub specific taxon 

(subspecies or population); appended to the 
global rank for the full species. The S Rank 
following applies to the subspecific taxon. 

? Denotes inexactness or uncertainty. 



© 2004 Montana Natural Heritage Program 



HowTO Read the Lists 



Additional Information 



Species are listed alphabetically by common name 
within major groups, which are organized alphabetically 
under Vertebrates and Invertebrates. Species with names 
bolded and underlined are new additions to the Species of 
Concern or Potential Species of Concern list. (Those 
downgraded from Concern to Potential Concern are not 
shown as new.) Where the global or state rank was 
changed, that rank is bolded and underlined. 

Current federal agency status designations are also 
provided, including legal status under the U.S. 
Endangered Species Act, and administrative designations 
of the U.S. Forest Service and Bureau of Land 
Management. Where the ESA listing status has changed 
since 2003, the new status is bolded and underlined. 

Federal Status Designations 

Fish AND Wildlife Service 

This column indicates status under the federal Endangered 
Species Act of 1973 based on categories defined by the 
U.S. Fish and Wildlife Service (16 U.S.C.A. § 1531-1543 
(Supp. 1996)). 

E Listed Endangered 

T Listed Threatened 

PE Proposed Endangered 

PT Proposed Threatened 

XN Experimental Nonessential 

C Candidate (species for which the U.S. Fish and 
Wildlife Service has sufficient information on 
biological status and threats to propose listing as 
threatened or endangered) 

Forest Service 

The U.S. Forest Service Manual (2670.22) defines the 
status of Sensitive species on Forest Service lands. The 
Regional Forester (Northern Region) designates Sensitive 
species on National Forests in Montana. This designation 
applies only on USFS-administered lands. 

S Sensitive; animal species identified by the 

Regional Forester for which population viability is 
a concern as evidenced by significant downward 
trend in population or a significant downward 
trend in habitat capacity. 

Bureau of Land Management 

The BLM 6840 Manual defines the status of species on 
Bureau of Land Management lands. The Montana State 
Office of the BLM assigns these designations in 1996. 
They apply only on BLM- administered lands. 

S Sensitive; species that are proven imperiled in at 

least part of their ranges and are documented to 
occur on BLM lands. 

© 2004 Montana Natural Heritage Program 



Our internet-based Field Guide to Montana's Animals, 

completed in 2003, provides information on over 600 
vertebrate species, including Species of Concern and 
Potential Species of Concern. The Field Guide offers a broad 
range of information, including species identification, range, 
habitat, reproduction, food habits, ecology, management, 
and references. For many Species of Concern, the Field 
Guide provides extensive detailed information from the 
databases of MTNHR Photos and call recordings are 
provided where available. 

The Field Guide can be accessed through the Natural 
Heritage Program web page at mtnhp.org and the Montana 
Fish, Wildlife and Parks web page at www.fwp.state.mt.us . 
The MTNHP website also offers the Montana Bird 
Distribution Database, where anyone can enter - and 
access - statewide bird observation data. 

The NatureServe Explorer website provides information 
on the status of species throughout North America. It can 
be found at w w w. natureser ve . org/explorer or from a link on 
the MTNHP website. This application includes status maps 
showing the state/provincial ranks across each species' 
range, assembled from the network of natural heritage 
programs. (Note that updated ranks for Montana animals 
will not be immediately reflected in NatureServe Explorer.) 

More detailed data on the occurrence of Montana 
Species of Concern are available on request from MTNHP, 
using the "Get Data" function on the website or through the 
contact information provided there. 

This report is available in electronic format on the 
MTNHP website under "Reports." Hard copies are 
available on request from MTNHP (see box below) or from 
MFWP at (406) 444-2535. 

Comments and Feedback 

We welcome comments and suggestions, including 
recommendations for additions or insights into species' 
status, as well as new observation data. All comments and 
information should be directed to MTNHP at 
mtnhp @ state. mt.us or the address below. Please submit 
observations using forms available on the MTNHP website 
(select "Animals" then "Submit Data"). 



Montana Natural Heritage Program 

P.O. Box 201800 
1515 East Sixth Avenue 
Helena, MT 59620-1800 
(406) 444-5354 Fax (406) 444-058 1 
mtnhp.org 
mtnhp @ state. mt.us 



u 



Selected References 

Brown, C.J. D. 1971. Fishes of Montana. Montana State 
University, Bozeman. 207 pp. 

Carlson, J. C. 2001. MontanaAnimal Species of Concern. 
Montana Natural Heritage Program and Montana 
Fish, Wildlife and Parks, Helena, Montana. 

Carlson, J. C. 2003. MontanaAnimal Species of Concern. 
Montana Natural Heritage Program and Montana 
Fish, Wildlife and Parks, Helen, Montana. 

Flath, D. L. 1984. Vertebrate species of special interest or 
concern. Montana Department of Fish, Wildlife and 
Parks. Helena. 76 pp. 

Flath, D. L. 1998. Species of special interest or concern. 
Montana Department of Fish, Wildlife and Parks, 
Helena, Montana. 7 pp. 

Frest, T. J. and E. J. Johannes. 1995. Interior Columbia 
Basin mollusk species of special concern. Final 
report to the Interior Columbia Basin Ecosystem 
Management Project, Walla Walla, WA. 274 pp. plus 
appendices. 

Foresman, K.R. 2001. The Wild Mammals of Montana. 
Special Publication No. 12. The American Society of 
Mammalogists. Lawrence, KS. 278 pp. 

Holton, G. D., and H. E. Johnson. 1996. Afield guide to 
Montana fishes. Second Edition. Mont. Dept. Fish, 
Wildl. Parks, Helena. 104 pp. 

Lenard, S., J. Carlson, J. Ellis, C. Jones, and C. Tilly. 2003. 
P.D. Skaar's Montana bird distribution. Sixth edition. 
Montana Audubon, Helena, MT. 144 pp. 

Lewis, J. J. 2001. Three new species of subterranean 
assellids from western North America, with a 
synopsis of the species of the region (Crustacea: 
Isopoda: Asellidae). Texas Memorial Museum, 
Speleological Monograpsh 5:1-15. 

Masters, L.L., L.E. Morse, A. S. Weakley, G.A. Hammerson 
andD.Faber-Langendoen. 2003. NatureServe 
Conservation Status Assessment Criteria. 
NatureServe, Arlington, VA, USA. 

Maxell, B. A., J. K. Werner, R Hendricks, and D. L. Flath. 
2003. Herpetology in Montana: a history, status 
summary, checklists, dichotomous keys, accounts for 
native, potentially native, and exotic species, and 
indexed bibliography. Northwest Fauna Number 5. 
138 pp. 



w 



Miller, K. B. and D. L. Gustafson. 1996. Distribution of the 
Odonata of Montana. Bulletin of American 
Odonatology3(4):75-88. 

Montana Bird Distribution Committee. 1996. P.D. Skaar's 
Montana bird distribution. Fifth edition. Special 
Publication No. 3, Montana Natural Heritage Program, 
Helena. 129 pp. 

Roedel, M. D. 1999. Montana animal species of special 
concern. Montana Natural Heritage Program, Helena. 
8 pp. 

Reichel,J.D.andD.Flath. 1995. Identification of 
Montana's amphibians and reptiles. Montana 
Outdoors 26(3): 15-34. 

Thompson, L. S. 1982. Distribution of Montana amphibians, 
reptiles, and mammals. Montana Audubon Council, 
Helena. 24 pp. 

Wang, D., and J. R. Holsinger. 2001. Systematics of the 
subterranean amphipod genus Stygobromus 
(Crangonyctidae) in western North America, with 
emphasis on the hubbsi group. Amphipacifica 3:39- 
147. 

Wright, PL. 1996. Statusof Rare Birds in Montana with 
Comments on Known Hybrids. Northwest Naturalist 

77(3):57-85. 



The preferred citation for this document is: 

Montana Natural Heritage Program. 2004. Montana 
Animal Species of Concern. MTNHP & MFWP. 
Helena, Montana. 11pp. 



Acknowledgements 

A team of MTNHP and MFWP staff coordinated the 
rank review process and production of this report; they 
include Steve Carson, Sue Crispin, Coburn Currier, Paul 
Hendricks, Janet Hess-Herbert, Andrew Jakes, Susan Lenard, 
Adam Messer, T.O. Smith and Leni Wilsmann. Thanks also 
to John Carlson who, as former Zoology Program Manager 
for MTNHP, got the ball rolling and proposed the ranking 
methodology now in use. 

We would also like to recognize and extend special 
thanks to the many, many biologists who have provided data 
and information, conducted status evaluations and reviewed 
species ranks throughout this process. 



© 2004 Montana Natural Heritage Program 



Common Name 



Species OF Concern 

Global 
Scientific Name Rank 



State 

Rank USFWS 



USPS BLM 



Amphibians 


Boreal Toad 


Bufo boreas boreas 


G4T4 


S2 






Coeurd'Alene Salamander 


Plethodon idahoensis 


G4 


S2 


S 


S 


Great Plains Toad 


Bufo cognatus 


G5 


S2 






Northern Leopard Frog 


Rana pipiens (E. of cont. divide) 


G5 


S3 


S 




Northern Leopard Frog 


Rana pipiens (W. of cont. divide) 


G5 


S1 


S 




Plains Spadefoot 


Spea bombifrons 


G5 


S3 






Birds 


Alder Flycatcher 


Empidonax ainorum 


G5 


S1B 






American White Pelican 


Pelecanus erytlirorliynclios 


G3 


S3B 






Baird's Sparrow 


Ammodramus bairdii 


G4 


S2B 


S 


S 


Bald Eagle 


Haliaeetus leucoceplialus 


G4 


S3B,S3N T 






Barn Owl 


Tyto alba 


G5 


S1B,S1N 






Black Rosy-Finch 


Leucosticte atrata 


G4 


S2 






Black Swift 


Cypseloides niger 


G4 


S3B 






Black Tern 


Ctilidonias niger 


G4 


S3B 




S 


Black-and-white Warbler 


Mniotilta varia 


G5 


S2S3B 






Black-backed Woodpecker 


Picoides arcticus 


G5 


S2 


S 


S 


Black-crowned Night-heron 


Nycticorax nycticorax 


G5 


S3B 






Blue-gray Gnatcatcher 


Polioptila caerulea 


G5 


S1B 






Bobolink 


Doliciionyx oryzivorus 


G5 


S2B 






Boreal Chickadee 


Poecile tiudsonica 


G5 


S1S2 






Brewer's Sparrow 


Spizella brewer i 


G5 


S2B 






Broad-tailed Hummingbird 


Selaspliorus platycercus 


G5 


S1B 






Burrowing Owl 


AWene cunicularia 


G4 


S2B 


S 


S 


Caspian Tern 


Sterna caspia 


G5 


S2B 






Cassin's Kingbird 


Tyrannus vociferans 


G5 


S2B 






Chestnut-collared LonasDur 


Calcarius ornatus 


G5 


S3B 






Columbian Sharp-tailed Grouse 


Tympanuctius ptiasianellus columbianus G4T3 


S1 


S 


S 


Common Loon 


Gavia immer 


G5 


S2B 


S 


s 


Common Tern 


Sterna iiirundo 


G5 


S3B 






Dickcissel 


Spiza americana 


G5 


S1S2B 




s 


Eastern Bluebird 


Sialia sialis 


G5 


S2B 






Ferruginous Hawk 


Buteo regalis 


G4 


S2B 




s 


Flammulated Owl 


Otus flammeolus 


G4 


S3B 


S 


s 


Forster's Tern 


Sterna forsteri 


G5 


S2B 






Franklin's Gull 


Larus pipixcan 


G4G5 


S3B 






Grasshopper Sparrow 


Ammodramus savannarum 


G5 


S3B 






Gray-crowned Rosy-Finch 


Leucosticte tepiirocotis 


G5 


S2B.S5N 






Great Gray Owl 


Strix nebuiosa 


G5 


S3 




s 


Greater Saae-Grouse 


Centrocercus uroplnasianus 


G4 


S3 


S 


s 


Harlequin Duck 


Histrionicus liistrionicus 


G4 


S2B 


S 


s 


Interior Least Tern 


Sterna antiiiarum atiiaiassos 


G4T2Q 


S1B E 






Lark Buntinq 


Caiamospiza meianocorys 


G5 


S3B 






Le Conte's Sparrow 


Ammodramus ieconteii 


G4 


S1S2B 




s 


Lewis's Woodpecker 


Meianerpes iewis 


G4 


S2B 






Loaaerhead Shrike 


Lanius iudovicianus 


G4 


S3B 


S 


s 


Lonq-billed Curlew 


Numenius americanus 


G5 


S2B 




s 


McCown's Lonqspur 


Caicarius mccownii 


G5 


S2B 






Mountain Plover 


CInaradrius montanus 


G2 


S2B 







© 2004 Montana Natural Heritage Program 



Common Name 



Scientific Name 



Global 
Rank 



State 

Rank USFWS 



USPS 



BLM 



Birds (con't) 


Nelson's Sharp-tailed Sparrow 


Ammodramus nelsoni 


G5 


S1B 








Northern Goshawk 


Accipiter gent His 


G5 


S3 




S 


s 


Northern Hawk Owl 


Surnia ulula 


G5 


S1B,S1N 








Olive-sided Flycatcher 


Contopus cooperi 


G4 


S3B 








Peregrine Falcon 


Faico peregrinus 


G4 


S2B 




S 


s 


Piping Plover 


Charadrius melodus 


G3 


S2B 


T 






Red-headed Woodpecker 


Melanerpes erythrocephalus 


G5 


S3B 








Saqe Sparrow 


Amphispiza belli 


G5 


S1S3B 






s 


Saae Thrasher 


Oreoscoptes montanus 


G5 


S3B 








Sedge Wren 


Cistothorus platensis 


G5 


S1B 








Sprague's Pipit 


A nth us spragueii 


G4 


S2B 




s 




Swainson's Hawk 


Buteo swainsoni 


G5 


S3B 






s 


Trumpeter Swan 


Cygnus buccinator 


G4 


S2B,S2N 




s 


s 


White-faced Ibis 


Plegadis chihi 


G5 


S1B 






s 


White-tailed Ptarmigan 


Lagopus leucurus 


G5 


S3 








Whooping Crane 


Grus americana 


G1 


S1M 


E 






Yellow Rail 


Coturn leaps noveboracensis 


G4 


S1B 








Yellow-billed Cuckoo 


Coccyzus americanus (E. pop.) 


G5 


S3B 








Yellow-billed Cuckoo 


Coccyzus americanus (W. pop.) 


G5 


S1S2B 


C 






Fish 


Blue Sucker 


Cycleptus elongatus 


G3G4 


S2S3 






s 


Bull Trout 


Salvelinus confluentus 


G3 


S2 


T 


s 




Columbia River Redband Trout 


Oncorhynchus mykiss gairdneri 


G5T4 


S1 




s 




Fluvial Arctic Grayling 


Thymallus arcticus montanus 


G5T1Q 


S1 


C 


s 




Northern Redbelly X Finescale Dace 


Phoxinus eos x phoxinus neogaeus 


HYB 


S3 






s 


Paddlefish 


Polyodon spathula 


G4 


S1S2 






s 


Pallid Sturgeon 


Scaphirhynchus albus 


G1 


S1 


E 






Pearl Dace 


Margariscus margarita 


G5 


S2 






s 


Sauger 


Stizostedion canadense 


G5 


S2 








Shortnose Gar 


Lepisosteus platostomus 


G5 


S1 






s 


Sicklefin Chub 


Macrohybopsis meeki 


G3 


S1 




s 


s 


Spoonhead Sculpin 


Cottus ricei 


G5 


S3 








Sturgeon Chub 


Macrohybopsis gelida 


G3 


S2 






s 


Torrent Sculpin 


Cottus rhotheus 


G5 


S3 




s 




Trout-perch 


Percopsis omiscomaycus 


G5 


S2 








Westslope Cutthroat Trout 


Oncorhynchus clarki lewis i 


G4T3 


S2 




s 


s 


White Sturgeon 


Acipenser transmontanus 


G4T1Q 


S1 


E 


s 




(Kootenai River Population) 














Yellowstone Cutthroat Trout 


Oncorhynchus clarki bouvieri 


G4T2 


S2 




s 


s 


l\/lammals 


American Bison 


Bos bison 


G4 


S2 








(free ranging herds) 














Arctic Shrew 


Sorex arcticus 


G5 


S1S3 








Black-footed Ferret 


Mustela nigripes 


G1 


S1 


E,XN 






Black-tailed Jack Rabbit 


Lepus californicus 


G5 


S2 








Black-tailed Prairie Dog 


Cynomys ludovicianus 


G4 


S3 


C 


s 


s 


Canada Lynx 


Lynx canadensis 


G5 


S3 


T 






Dwarf Shrew 


Sorex nanus 


G4 


S2S3 








Eastern Red Bat 


Lasiurus borealis 


G5 


S2S3 








Fisher 


Martes pennanti 


G5 


S3 




s 


s 



© 2004 Montana Natural Heritage Program 







Global 


State 




Common Name 


Scientific Name 


Rank 


Rank USFWS 


USPS BLM 


Mammals (con't) 


Fringed Myotis 


Myotis thysanodes 


G4G5 


S3 




Gray Wolf 


Canis lupus 


G4 


S3 T,XN 




Grizzly Bear 


Ursus arctos horribilis 


G4T3T4 


S2S3 T 




Great Basin Pocket Mouse 


Perognathus parvus 


G5 


S2S3 




Hispid Pocket Mouse 


Chaetodipus hispidus 


G5 


S1S3 




Meadow Jumping Mouse 


Zapus hudsonius 


G5 


S2 


s 


Merriam's Shrew 


Sorex merriami 


G5 


S3 


s 


North American Wolverine 


Gulo gulo luscus 


G4T4 


S3 


s s 


Northern Bog Lemming 


Synaptomys borealis 


G4 


S2 


s s 


Northern Myotis 


Myotis septentrionalis 


G4 


S2S3 




Pallid Bat 


Antrozous pallidus 


G5 


S2 




Preble's Shrew 


Sorex preblei 


G4 


S3 


s 


Pygmy Rabbit 


Brachylagus idahoensis 


G4 


S3 


s s 


Spotted Bat 


Euderma maculatum 


G4 


S2 




Swift Fox 


Vulpes velox 


G3 


S3 


s s 


Townsend's Big-eared Bat 


Corynorhinus townsendii 


G4 


S2 


s s 


Uinta Chipmunk 


Tamiasumbrinus 


G5 


S3 




White-tailed Prairie Dog 


Cynomys leucurus 


G4 


S1 


s s 


Reptiles 


Greater Short-horned Lizard 


Plirynosoma hernandesi 


G5 


S3 




Milk Snake 


Lampropeltis triangulum 


G5 


S2 




Northern Alligator Lizard 


Elgaria coerulea 


G5 


S3 




Sagebrush Lizard 


Sceloporus graciosus 


G5 


S3 




Smooth Green Snake 


Liochlorophis vernalis 


G5 


Si 




Snapping Turtle 


Chelydra serpentina 


G5 


S3 


s 


Spiny Softshell 


Apalone spinifera 


G5 


S3 


s 


Western Hognose Snake 


Heterodon nasicus 


G5 


S2 




Western Skink 


Eumeces skiltonianus 


G5 


S3 




Invertebrates - Insects 


Beetles 










Brown's Microcylloepus Riffle Beetle 


Microcylloepus brown i 


G1 


S1 




Warm Spring Zaitzevian Riffle Beetle 


Zaitzevia thermae 


G1 


S1 C 




Butterflies 










Alberta Fritillarv 


Boloria alberta 


G3 


S2S3 




Friaaa's Fritillarv 


Boloria frigga 


G5 


S1S2 




Gillette's Checkerspot 


Euphydryas gillettii 


G2G3 


S2 




Grav Comma 


Polygonia progne 


G5 


S2 




Maqdalena Alpine 


Erebia magdalena 


G5 


S2 




Ottoe Skipper 


Hesperia ottoe 


G3G4 


S2S3 


s 


Caddlsflles 










A Caddisfly 


Rhyacophila ebria 


G1? 


S1? 




A Rhyacophilan Caddisfly 


Rhyacophila newel li 


G2? 


S2? 




A Rhyacophilan Caddisfly 


Rhyacophila glacieri 


G1 


S1 




Alexander's Rhyacophilan Caddisfly 


Rhyacophila alexanderi 


G2 


S2 




An Agapetus Caddisfly 


Agapetus montanus 


G2? 


S2? 




Damselflles 










Last Best Place Damselfly 


Enallagma optimolocus 


G1G3Q 


S1S3 




Subarctic Bluet 


Coenagrion interrogatum 


G5 


S1S2 




Dragonflles 










Boreal Whiteface 


Leucorrhinia borealis 


G5 


S1 





© 2004 Montana Natural Heritage Program 



Common Name 



Scientific Name 



Global 
Rank 



State 

Rank USFWS 



USPS BLM 



Invertebrates - Insects (con't) 


Dragon flies (con't) 








Brush-tipped Emerald 


Somatochlora walshii 


G5 


S1S2 


Eastern Ringtail 


Erpetogomphus designatus 


G5 


S1 


Subarctic Darner 


Aeshna subarctica 


G5 


S1S2 


Western Pondhawk 


Erythemis collocata 


G5 


S1S2 


Mayflies 








A Mayfly 


Caenis youngi 


G4 


S2 


Sprlngtalis 








ASpringtail 


Oncopodura cruciata 


G1G2 


S1S2 


Stoneflles 








A Stonefly 


Isocapnia crinita 


G4 


S2 


A Stonefly 


Isocapnia Integra 


G4 


S2 


A Stonefly 


Utacapnia columbiana 


G4 


S2 


A Stonefly 


Zapada cordillera 


G3 


S2 


A Stonefly 


Isoperia petersoni 


G5 


S2 


Meltwater Lednian Stonefly 


Lednia tumana 


G1 


S1 


Western Glacier Stonefly 


Zapada glacier 


G2 


S1 


Invertebrates - Mollusks 


Alpine Mountainsnail 


Oreohelix alpina 


G1 


S1 


Berry's Mountainsnail 


Oreohelix strigosa berryi 


G5T2 


S1S2 


Bitterroot Mountainsnail 


Oreohelix amariradix 


G1G2 


S1S2 


Carinate Mountainsnail 


Oreohelix eirodi 


G1 


S1 


Fir Pinwheel 


Radiodiscus abietum 


G3 


S2S3 


Flathead Pondsnail 


Stagnicola eirodi 


G1 


S1 


Gallatin Mountainsnail 


Oreohelix yavapai mariae 


G4T1Q 


S1 


Humoed Coin 


Polygyrella polygyrella 


G2G3 


S1S3 


Keeled Mountainsnail 


Oreohelix carinifera 


G1 


S1 


Large-mantle Physa 


Physa megalochlamys 


G3 


S1 


Largemouth Pondsnail 


Stagnicola eirodiana 


G1 


S1 


Lyre IVIantlesluq 


Udosarx lyrata lyrata 


G2T2 


SI 


Marbled Jumping-slug 


Hemphillia danieisi 


G1G3 


S1S3 


Mission Range Disc 


Discus brunsoni 


G1 


S1 


Mountain Marshsnail 


Stagnicola montanensis 


G3 


S1S3 


Rocky Mountain Capshell 


Acroloxus color adensis 


G1G2 


S1 


Rocky Mountain Duskysnail 


Colligyrus greggi 


G3G4 


S1 


Russell Mantleslug 


Udosarx lyrata russelli 


G2T1 


S1 


Sheathed Slug 


Zacoleus idahoensis 


G3G4 


S2S3 


Shortface Lanx 


Fisherola nuttalli 


G2 


S1S3 


Spotted Slug 


Magnipelta mycophaga 


G2G3 


S1S3 


Striate Disc 


Discus shimekii 


G5 


S1 


Invertebrates - Other 


Arachnids 








A cave obligate Harvestman 


Cryptobunus cavicolus 


G1G2 


S1S2 


Crustaceans 








A cave obligate Amphipod 


Stygobromus glacialis 


G1G2 


S1S2 


A cave obligate Amphipod 


Stygobromus montanensis 


G1G2 


S1S2 


A cave obligate Amphipod 


Stygobromus obscurus 


G1G2 


S1S2 


A cave obligate Amphipod 


Stygobromus puteanus 


G1G2 


S1S2 


A cave obligate Amphipod 


Stygobromus tritus 


G1G2 


S1S2 


A cave obligate Isopod 


Salmasellus steganothrix 


G2G3 


S1S2 



© 2004 Montana Natural Heritage Program 



Potential Species of Concern 



Common Name 



Scientific Name 



Global 
Rank 



State 
Rank 



USFWS USPS 



BLM 



Birds 


American Three-toed Woodpecker 


Picoides dorsalis 


G5 


S3S4 




Baltimore Oriole 


Icterus galbula 


G5 


S3S4B 




Black-billed Cuckoo 


Coccyzus erythropthalmus 


G5 


S3S4B 




Black-necked Stilt 


Himantopus mexicanus 


G5 


S3S4B 




Chimney Swift 


Chaetura pelagica 


G5 


S3S4B 




Clark's Grebe 


Aechmophorus clarkii 


G5 


S2S4B 




Common Poorwill 


Phalaenoptilus nuttallii 


G5 


S3S4B 




Eastern Screech-Owl 


Megascops asio 


G5 


S3S4 




Great Blue Heron 


Ardea herodias 


G5 


S3S4 




Horned Grebe 


Podiceps auritus 


G5 


S4 




Indiqo Buntinq 


Passerina cyanea 


G5 


S2S4B 




Orchard Oriole 


Icterus spurius 


G5 


S2S4B 




Ovenbird 


Seiurus aurocapilla 


G5 


S3S4B 




Plumbeous Vireo 


Vireo plumbeus 


G5 


S3S4B 




Short-eared Owl 


Asio flammeus 


G5 


S3S4 




Tennessee Warbler 


Vermivora peregrina 


G5 


S2S4B 




Western Screech-Owl 


Megascops kennicottii 


G5 


S3S4 




Williamson's Sapsucker 


Sphyrapicus thyroideus 


G5 


S3S4B 




Fish 


Brassy Minnow 


Hybognathus hankinsoni 


G5 


su 




Brook Stickleback 


Culaea inconstans 


G5 


su 




Burbot 


Lota lota 


G5 


su 


S 


Creek Chub 


Semotilus atromaculatus 


G5 


su 




Iowa Darter 


Etheostoma exile 


G5 


su 




Lake Trout (Native Population) 


Salvelinus namaycush 


G5 


su 




Northern Red-bellied Dace 


Phoxinus eos 


G5 


su 




Plains Minnow 


Hybognathus placitus 


G4 


su 




Pygmy Whitefish 


Prosopium coulter! 


G5 


su 




Shorthead Sculpin 


Cottus confusus 


G5 


su 




l\/lammals 


Hayden's Shrew 


Sorex haydeni 


G4 


S3S4 




Hoary Bat 


Lasiurus cinereus 


G5 


S3S4 




Hoarv Marmot 


Marmota calagata 


G5 


S3S4 




Idaho Pocket Gopher 


Thomomys idahoensis 


G4 


S2S4 




Silver-haired Bat 


Lasionycteris noctivagans 


G5 


S3S4 




Uinta Ground Sauirrel 


Spermophilus armatus 


G5 


S3S4 




Western Spotted Skunk 


Spilogale gracilis 


G5 


S2S4 




Wvomina Ground Sauirrel 


Spermophilus elegans 


G5 


S3S4 




Yuma Myotis 


Myotis yumanensis 


G5 


S3S4 




Invertebrates - Insects 


Beetles 










A Tiqer Beetle 


Cicindela cuprascens 


G5 


S3 




A Tiqer Beetle 


Cicindela limbata 


G5 


S3 




Butterflies 










Astarte FritillarY 


Boloria astarte 


G5 


S2S3 




Colorado Alpine 


Erebia callias 


G5 


S2S3 




Damoetus Checkerspot 


Chlosyne damoetas 


G4G5 


S3S5 




Eyed Brown 


Satyrodes eurydice 


G4 


S2S3 





© 2004 Montana Natural Heritage Program 



Common Name 



Scientific Name 



Global 
Rank 



State 
Rank 



USFWS 



USPS BLM 



Invertebrates - Insects (co 


n't) 








Butterflies {con \) 










Freiia Fritillarv 


Boloria freija 


G5 


S3S5 




Giant Sulfur 


Colias gigantea 


G5 


S3 




Indra Swallowtail 


Papillo indra 


G5 


S2S3 




Labrador Sulfur 


Colias nastes 


G5 


S2S3 




Melissa Arctic 


Oeneis melissa 


05 


S2S3 




Mormon Metalmark 


Apodemia mormo 


G5 


S3S5 




Northern Marble 


Euchloe creusa 


G5 


S1S3 




Red-disked Alpine 


Erebia discoidalis 


05 


S3S5 




Tawny Crescent 


Phyciodes batesii 


G4 


S2S3 


S 


White Admiral 


Limenitis arthemis 


G5 


S2S3 




White-veined Arctic 


Oeneis bore 


05 


S2S3 




Damselflies 










Alkali Bluet 


Enallagma clausum 


G5 


S2S4 




Arrovo Bluet 


Enallagma prae varum 


G5 


S3S5 




Emma's Dancer 


Argia emma 


05 


S3S5 




Familiar Bluet 


Enallagma civile 


G5 


S2S4 




Paiute Dancer 


Argia alaberta 


G4 


S2S3 




Prairie Bluet 


Coenagrion angulatum 


G5 


S1S3 




Vivid Dancer 


Argia vidida 


G5 


S3S5 




Dragonflies 










Black-tipped Darner 


Aeshna tuberculifera 


G4 


S2S4 




Blue-eyed Darner 


Aeshna multicolor 


G5 


S2S4 




California Darner 


Aeshna calif or nica 


G5 


S3S5 




Chalk-fronted Skimmer 


Libellua Julia 


G5 


S3S4 




Common Whitetail 


Libellula lydia 


G5 


S3S5 




Crimson-winqed Whiteface 


Leucorrhinia glacialis 


G5 


S3 




Eiaht-SDOtted Skimmer 


Libellula forensis 


G5 


S3S5 




Flame Skimmer 


Libellula saturata 


G5 


S2S4 




Horned Clubtail 


Arigomphus cornutus 


G4 


S2S4 




Hudsonian Emerald 


Somatochlora hudsonica 


G5 


S2S4 




Lake Darner 


Aeshna eremita 


G5 


S3S4 




Lance-tipped Darner 


Aeshna constricta 


G5 


S1S3 




Mountain Emerald 


Somatochlora semicircularis 


G5 


S3S5 




Ocellated Emerald 


Somatochlora minor 


G5 


S2S4 




Plains Clubtail 


Gomphus externus 


G5 


S2S4 




Pronqhorn Clubtail 


Gomphus graslinellus 


G5 


S3S5 




Red-veined Meadowhawk 


Sympetrum madidum 


G4 


S2S3 




Ringed Emerald 


Somatochlora albicincta 


G5 


S1S3 




River Jewelwinq 


Calopteryz aequabilis 


G5 


S3S4 




Rush Darner 


Aeshna juncea 


G5 


S3S5 




Sinuous Snaketail 


Ophiogomphus occidentis 


G4 


S2S4 




SDinv Baskettail 


Epitheca spinigera 


G5 


S3S5 




Twelve-spotted Skimmer 


Libellula pulchella 


G5 


S3S5 




Zigzag Darner 


Aeshna sitchensis 


G5 


S2S3 




Invertebrates - Mollusks 


Threeridqe Valvata 


Valvata tricarnata 


G5 


S2S3 




Western Pearlshell 


Margaritifera falcata 


G4 


S2S4 





© 2004 Montana Natural Heritage Program 



10 



Species Removed from Concern 



Common Name 


Scientific Name 


Global 
Rank 


State 
Rank 


Mammals 


Pygmy Shrew 


Sorex hoyi 


G5 


S4 



Species Removed from Review 



Common Name 


Scientific Name 


Global 
Rank 


State 
Rank 


Biitls 


Cassin's Vireo 
Wilson's Phalarope 


Vireo cassinii 
Phalaropus tricolor 


G5 
G5 


S4 
S4 


Mammals 


Eastern Cottontail 


Sylvilagus floridanus 


G5 


S4 


Reptiles 


Common Garter Snake 


Thamnophis sirtalis 


G5 


S4 


Damselflies 


Sweetflag Spreadwing 
Variable Dancer 


Lestes forcipatus 

Argia Fumipennis violacea 


G5 
G5T5 


SU 
SU 


Dragonflies 


Canada Darner 
Pacific Spiketail 
Plains Emerald 
Wandering Glider 


Aeshna canadensis 
Cordulegaster hageni 
Somatochlora ensigera 
Pantala flavescens 


G5 
G5 
G4 
G5 


SU 
SU 
SU 
SU 


Mollusks 


Columbia Pebblesnail 
Prairie Sprite 


Fluminicola fuscus G3 
Promenetus exacuous megas G5TU 


SU 
SU 



© 2004 Montana Natural Heritage Program 



11